Explain mechanisms by which two homologous chromosomes are tied with each other in metaphase I (metaphase in the first meiosis) and by which this tie is lost at the onset of anaphase I (anaphase in...

Explain mechanisms by which two homologous chromosomes are tied with each other in metaphase I (metaphase in the first meiosis) and by which this tie is lost at the onset of anaphase I (anaphase in the first meiosis). In your answer include an explanation of the mechanism by which sister chromatids are held together during anaphase I.